Scapania bolanderi

Capsule valves vertical Medium: 1.5 - 2.5 mm wide
Dorsal base decurrent
Shoot anisophyllous
Leaf margin entire
Leaves spreading
Leaves divided >0.5
Leaf base not sheathing
Lobe tips rounded
Cell walls evenly thin
Vitta absent
Oil-bodies granulose
Perianth present
Perianth long emergent
Perianth flattened
Perianth mouth wide
Perianth mouth entire
Perigynium absent
Dioicous
Leaf bases not decurrent
Small: 0.5 - 1.5 mm wide
Shoot tips unreduced
Leaves closely imbricate
Leaves overlap slightly
Lobes folded together
Keel rounded
Both lobes decurrent
Underleaves absent
Trigones not thickened
Oil-bodies solitary
Oil-bodies brown
Gemmae round
Gemmae on leaf margins
No microphyllous branches
Leaf margin toothed
Leaf margin ciliate
Shoots mainly ascending
Large: 2.5 - 4 mm wide
Robust: > 4 mm wide
Dark, olive green
Green to reddish brown
Divided into 2 lobes
Keel sharp
Rhizoids scattered
Cell walls unevenly thick
Trigones concave
Trigones bulging
Cuticle smooth
Oil-bodies present
Oil-bodies ave 2-12/cell
Oil-bodies colorless
Perianth mouth dentate
Shoots mostly unbranched
Tinged with bright red
Dark brown to blackish
Leaves succubous
Leaves angled to stem
Leaves transverse
Ventral base decurrent
Ventral lobe larger
Dorsal lobe decurrent
Ventral lobe decurrent
Neither lobe decurrent
Lobe tips obtuse
Lobes triangular
Lobes rotund
Sinus acute
Rhizoids not prominent
Cell walls evenly thick
Marginal cells thicker
Cuticle striate/papillose
Oil-bodies ave 3-5/cell
Oil-bodies ave 5-8/cell
Oil-bodies clear
Perianth mouth ciliate
Gemmae present
Gemmae green
Gemmae red